Motivating students to engage with physical chemistry through biological examples, this textbook demonstrates how the tools of physical chemistry can be used to illuminate biological questions. It clearly explains key principles and their relevance to life science students, using only the most straightforward and relevant mathematical tools. More than 350 exercises are spread throughout the chapters, covering a wide range of biological applications and explaining issues that students often find challenging. Answers to the exercises are included in the book. Fully worked solutions and answers to the review problems, password-protected for instructors, are available at www.cambridge.org/roussel.